The town is on the A30 at its junction with the A323 Fleet Road, almost equidistant between Basingstoke to the west and Camberley to the east. This was the main trunk road to the West Country and Southampton from the stagecoach era until 1971, when the M3 motorway was opened. Its nearest junction is Junction 5, about 6 miles (10 km) southwest of the town.
At the Phœnix pub in 1934 a group of people formed the Vintage Sports-Car Club which has gone from strength to strength until it now numbers nearly 7,000 members with races and other sporting events all over the world.
The nearest railway station is at Winchfield on the South Western main line. It is about 1 1⁄2 miles (2.4 km) south of Hartley Wintney and is signposted from the town.
Stagecoach South and Reading Buses buses serve Hartley Wintney. Route 7 links Aldershot, Fleet and Reading via Hartley Wintney. In addition route 65X runs between Alton College and Phoenix Green via Hartley Wintney and route 408 runs between Farnborough College of Technology and Reading via Hartley Wintney.
Hartley Wintney Community Bus Service runs a regularly weekday commuter service to Winchfield railway station and services to Hook, Yateley, Frogmore and The Meadows. A Saturday service goes to Camberley as well as The Meadows from Hartley Wintney, via Yateley and Frogmore.
- Hook – 3 miles (5 km) west
- Fleet – 3 miles (5 km) southeast
- Yateley – 4 miles (6.4 km) northeast
- Sandhurst – 5 miles (8 km) northeast
- Farnborough – 6 miles (10 km) east-southeast
- Camberley – 7 miles (11 km) east-northeast
- Aldershot – 7 1⁄2 miles (12 km) southeast
- Basingstoke – 8 miles (13 km) west
- Reading – 11 miles (18 km) north
- London (central) – 39 miles (63 km) east-northeast