|Doctoral students||Gerald Guralnik|
Gilbert was born in
Gilbert returned to Harvard in 1957 and promoted in 1968 to professor of biochemistry.  In 1969, Gilbert was given Harvard's Ledlie Prize.  In 1972, Gilbert was named American Cancer Society Professor of Molecular Biology. 
He is a co-founder of the biotech start-up companies Biogen and Myriad Genetics. Gilbert is currently the chairman of the Harvard Society of Fellows.