Parrot

Parrots
Temporal range: EoceneHolocene 50–0 Ma
KakapoAustralian ringneckBlue-and-yellow macawKeaLesser vasa parrotPalm cockatooParrot montage.jpg
About this image
Scientific classification e
Kingdom:Animalia
Phylum:Chordata
Class:Aves
Clade:Psittacopasserae
Order:Psittaciformes
Wagler, 1830
Superfamilies

Cacatuoidea (cockatoos)
Psittacoidea (true parrots)
Strigopoidea (New Zealand parrots)

Parrot range.png
Range of parrots, all species (red)

Parrots, also known as psittacines z/,[1][2] are birds of the roughly 393 species in 92 genera that make up the order Psittaciformes, found in most tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoidea ("true" parrots), the Cacatuoidea (cockatoos), and the Strigopoidea (New Zealand parrots). Parrots have a generally pantropical distribution with several species inhabiting temperate regions in the Southern Hemisphere, as well. The greatest diversity of parrots is in South America and Australasia.

Characteristic features of parrots include a strong, curved bill, an upright stance, strong legs, and clawed zygodactyl feet. Many parrots are vividly coloured, and some are multi-coloured. Most parrots exhibit little or no sexual dimorphism in the visual spectrum. They form the most variably sized bird order in terms of length.The most important components of most parrots' diets are seeds, nuts, fruit, buds, and other plant material. A few species sometimes eat animals and carrion, while the lories and lorikeets are specialised for feeding on floral nectar and soft fruits. Almost all parrots nest in tree hollows (or nest boxes in captivity), and lay white eggs from which hatch altricial (helpless) young.

Parrots, along with ravens, crows, jays, and magpies, are among the most intelligent birds, and the ability of some species to imitate human voices enhances their popularity as pets. Trapping wild parrots for the pet trade, as well as hunting, habitat loss, and competition from invasive species, has diminished wild populations, with parrots being subjected to more exploitation than any other group of birds. Measures taken to conserve the habitats of some high-profile charismatic species have also protected many of the less charismatic species living in the same ecosystems.

Taxonomy

Origins and evolution

Fossil dentary specimen UCMP 143274 restored as a parrot (left) or an oviraptorosaur

Psittaciform diversity in South America and Australasia suggests that the order may have evolved in Gondwana, centred in Australasia.[3] The scarcity of parrots in the fossil record, however, presents difficulties in confirming the hypothesis, and there is currently a higher amount of fossil remains from the northern hemisphere in the early Cenozoic.[4] Molecular studies suggest that parrots evolved approximately 59 million years ago (Mya) (range 66–51 Mya) in Gondwana. The three major clades of Neotropical parrots originated about 50 Mya (range 57–41 Mya).[5]

A single 15 mm (0.6 in) fragment from a large lower bill (UCMP 143274), found in deposits from the Lance Creek Formation in Niobrara County, Wyoming, had been thought to be the oldest parrot fossil and is presumed to have originated from the Late Cretaceous period, which makes it about 70 million years old.[6] However, other studies suggest that this fossil is not from a bird, but from a caenagnathid oviraptorosaur (a non-avian dinosaur with a birdlike beak), as several details of the fossil used to support its identity as a parrot are not actually exclusive to parrots, and it is dissimilar to the earliest-known unequivocal parrot fossils.[7][8]

It is generally assumed that the Psittaciformes were present during the Cretaceous–Paleogene extinction event (K-Pg extinction), 66 mya. They were probably generalised arboreal birds, and did not have the specialised crushing bills of modern species.[4][9] Genomic analysis provides strong evidence that parrots are the sister group of passerines, forming the clade Psittacopasserae, which is the sister group of the falcons.[10]

The first uncontroversial parrot fossils date to tropical Eocene Europe around 50 mya. Initially, a neoavian named Mopsitta tanta, uncovered in Denmark's Early Eocene Fur Formation and dated to 54 mya, was assigned to the Psittaciformes. However, the rather nondescript bone is not unequivocally psittaciform, and it may rather belong to the ibis genus Rhynchaeites, whose fossil legs were found in the same deposits.[11]

Fossil skull of a presumed parrot relative from the Eocene Green River Formation in Wyoming

Several fairly complete skeletons of parrot-like birds have been found in England and Germany.[12] These are probably not transitional fossils between ancestral and modern parrots, but rather lineages that evolved parallel to true parrots and cockatoos:[13]

The earliest records of modern parrots date to around 23–20 mya.[15] The fossil record—mainly from Europe—consists of bones clearly recognisable as belonging to anatomically modern parrots.[16] The Southern Hemisphere contains no known parrot-like remains earlier than the Early Miocene around 20 mya.[15]

Phylogeny

Parrots
 
 

Psittacoidea Rose-ringed Parakeet (Male) I IMG 9141.jpg

 

Cacatuoidea Cacatua galerita -perching on branch -crest-8a-2c.jpg

 

Strigopoidea Kaka (Nestor meridionalis)- Wellington -NZ-8-2c.jpg

 

Other birds

Phylogenetic relationship between the three parrot superfamilies[3][17][18]

The Psittaciformes comprise three main lineages: Strigopoidea, Psittacoidea and Cacatuoidea.[19] The Strigopoidea were considered part of the Psittacoidea, but the former is now placed at the base of the parrot tree next to the remaining members of the Psittacoidea, as well as all members of the Cacatuoidea.[3][17][18] The Cacatuoidea are quite distinct, having a movable head crest, a different arrangement of the carotid arteries, a gall bladder, differences in the skull bones, and lack the Dyck texture feathers that—in the Psittacidae—scatter light to produce the vibrant colours of so many parrots. Colourful feathers with high levels of psittacofulvin resist the feather-degrading bacterium Bacillus licheniformis better than white ones.[20] Lorikeets were previously regarded as a third family, Loriidae,[21]:45 but are now considered a tribe (Loriini) within the subfamily Loriinae, family Psittaculidae. The two other tribes in the subfamily are the closely related fig parrots (two genera in the tribe Cyclopsittini) and budgerigar (tribe Melopsittacini).[3][17][18]

 Strigopoidea 

Nestoridae 

Strigopidae 

 
 Cacatuoidea 

 

 Psittacoidea 
Psittacidae
 

Neotropical parrots

 

Psittacinae

Psittrichasiidae
 

Psittrichadinae

 

Coracopseinae

Psittaculidae

Psittaculinae

Broad-tailed parrots

Fig parrots

Budgerigar

Lories and Lorikeets

Bolbopsittacus

Hanging parrots

Lovebirds

Psittacella

Phylogenetic relations between parrots[3]

Systematics

The order Psittaciformes consists of roughly 393 species belonging to 92 genera.[22][3][17][19][23][24][25][26]

Skeleton of a parrot

Superfamily Strigopoidea: New Zealand parrots

Superfamily Cacatuoidea: cockatoos

Superfamily Psittacoidea: true parrots

Other Languages
Afrikaans: Papegaaie
Alemannisch: Papagei
አማርኛ: ብቅበቃ
Ænglisc: Bleohfugol
العربية: ببغاء
asturianu: Psittaciformes
Avañe'ẽ: Gua'a
azərbaycanca: Tutuquşukimilər
বাংলা: টিয়া
Bân-lâm-gú: Eng-ko
беларуская: Папугаепадобныя
беларуская (тарашкевіца)‎: Папугаепадобныя
भोजपुरी: सुग्गा
Bislama: Barod
български: Папагалоподобни
bosanski: Papiga
brezhoneg: Psittaciformes
буряад: Тоти
català: Psitaciformes
čeština: Papoušci
Cymraeg: Parot
dansk: Papegøje
Deutsch: Papageien
Diné bizaad: Tsídii yáłtiʼí
डोटेली: सुवा
Ελληνικά: Παπαγάλος
español: Psittaciformes
Esperanto: Papagoformaj
eʋegbe: Ako
فارسی: طوطی
føroyskt: Papageykar
français: Psittaciformes
Gaeilge: Pearóid
Gàidhlig: Piorraid
ગુજરાતી: પોપટ
한국어: 앵무새
हिन्दी: तोता
hrvatski: Papigašice
Bahasa Indonesia: Bayan (burung)
interlingua: Psittaciformes
íslenska: Páfagaukar
italiano: Psittaciformes
עברית: תוכאים
Jawa: Bèthèt
ಕನ್ನಡ: ಗಿಳಿ
Kiswahili: Kasuku
Kreyòl ayisyen: Jako
kurdî: Tûtî
Кыргызча: Тотулар
кырык мары: Попугай
latviešu: Papagaiļi
Lëtzebuergesch: Papageien
Ligure: Pappagaggio
Limburgs: Pappegejje
Lingua Franca Nova: Psitasiformo
lumbaart: Psittaciformes
македонски: Папагаловидни
Bahasa Melayu: Psittaciformes
Mìng-dĕ̤ng-ngṳ̄: Ĕng-gŏ̤
мокшень: Попугай
монгол: Тоть
မြန်မာဘာသာ: ကြက်တူရွေး
Nederlands: Papegaaiachtigen
नेपाली: सुगा
日本語: オウム目
Nordfriisk: Papagein
norsk nynorsk: Papegøyefuglar
ଓଡ଼ିଆ: ଶୁଆ
oʻzbekcha/ўзбекча: Toʻtilar
ਪੰਜਾਬੀ: ਤੋਤਾ
پنجابی: طوطا
پښتو: توتي
Patois: Paarat
Перем Коми: Попугай
Plattdüütsch: Papagoyen
polski: Papugowe
português: Psittaciformes
română: Psittaciformes
ᱥᱟᱱᱛᱟᱲᱤ: ᱢᱤᱨᱩ
संस्कृतम्: शुकः
Scots: Paurit
shqip: Papagalli
Simple English: Parrot
سنڌي: طوطو
slovenčina: Papagájotvaré
slovenščina: Papige
српски / srpski: Папагаји
srpskohrvatski / српскохрватски: Papigašice
Tagalog: Pikoy
தமிழ்: கிளி
తెలుగు: చిలుక
Türkçe: Papağansılar
удмурт: Попугай
українська: Папугоподібні
اردو: طوطا
Vahcuengh: Roegyengj
Tiếng Việt: Bộ Vẹt
West-Vlams: Poapegoais
Winaray: Pikóy
吴语: 鹦形目
ייִדיש: פאפוגיי
Yorùbá: Odídẹrẹ́
粵語: 鸚鵡
Zazaki: Papağan
中文: 鹦形目