Approximate area of the origin of L0d and L0k haplogroups in southern Africa, dated to before 90,000 years ago by Behar et al. (2008).
The ancestors of the modern Khoisan expanded to Southern Africa before 150,000 years ago, possibly as early as before 260,000 years ago, so that by the beginning of the MIS 5 "megadrought", 130,000 years ago, there were two ancestral population clusters in Africa, bearers of mt-DNA haplogroup L0 in southern Africa, ancestral to the Khoi-San, and bearers of haplogroup L1-6 in central/eastern Africa, ancestral to everyone else.
Due to their early expansion and separation, the populations ancestral to the Khoisan have been estimated as having represented the "largest human population" during the majority of the anatomically modern human timeline, from their early separation before 150 ka until the recent peopling of Eurasia some 70 kya.
They were much more widespread than today, their modern distribution being due to their decimation in the course of the Bantu expansion. They were dispersed throughout much of Southern and South-Eastern Africa.
There was also a significant back-migration of bearers of L0 towards eastern Africa between 120 and 75 kya.
Rito et al. (2013) speculate that pressure from such back-migration may even have contributed to the dispersal of East African populations out of Africa at about 70 kya.
"By ~130 ka two distinct groups of anatomically modern humans co-existed in Africa: broadly, the ancestors of many modern-day Khoe and San populations in the south and a second central/eastern African group that includes the ancestors of most extant worldwide populations. Early modern human dispersals correlate with climate changes, particularly the tropical African “megadroughts” of MIS 5 (marine isotope stage 5, 135–75 ka) which paradoxically may have facilitated expansions in central and eastern Africa, ultimately triggering the dispersal out of Africa of people carrying haplogroup L3 ~60 ka. Two south to east migrations are discernible within haplogroup L0. One, between 120 and 75 ka, represents the first unambiguous long-range modern human dispersal detected by mtDNA and might have allowed the dispersal of several markers of modernity. A second one, within the last 20 ka signalled by L0d, may have been responsible for the spread of southern click-consonant languages to eastern Africa, contrary to the view that these eastern examples constitute relics of an ancient, much wider distribution."
Approximate distribution of Middle Stone Age
Capoids at the end of the Pleistocene (12,000 years ago) shown in magenta (according to C. S. Coon
The term Capoid was introduced as a racial category uniting the Khoikhoi and San peoples as separate from
other sub-Saharan populations was proposed by Carleton S. Coon (1962). It is named for the Cape of Good Hope.
Coon (1962) proposed that the term "Negroid" should be abandoned, and the sub-Saharan African populations of West African stock (including the Bantu) should be termed "Congoid" instead.
Cavalli-Sforza at al. (1994) used the term Khoisanid instead of Coon's "Capoid".
The observation of a significant difference between the Khoisan and the populations of West African stock was not original to Coon, it had been noted as early as 1684 François Bernier, the early modern author who originally introduced the French word race to refer to the large divisions of mankind. Bernier outside of five large divisions described in more detail proposes the possible addition of more categories, primarily for "the Blacks of the Cape of Good Hope" (les Noirs du Cap de bonne Esperance), which seemed to him to be of significantly different build from most other populations below the Sahara.
Late Stone Age
Schematic representation of the "out of South Africa" migration of the post-Eemian Middle to Late Stone Age (after 100 kya) inferred from mtDNA haplogroup L0 in modern African populations (Rito et al. 2013).
The Khoisanid populations ancestral to the Khoisan were spread throughout much of Southern and Eastern Africa throughout the Late Stone Age, after about 75 ka. A further expansion, dated to about 20 ka, has been proposed based
on the distribution of the L0d haplogroup. Rosti et al. suggest a connection of this recent expansion with the spread of click consonants to eastern African languages (Hadza language).
The Late Stone Age Sangoan industry occupied southern Africa in areas where annual rainfall is less than a metre (1000 mm; 39.4 in). The contemporary San and Khoi peoples resemble those represented by the ancient Sangoan skeletal remains.
Against the traditional interpretation that finds a common origin for the Khoi and San, other evidence has suggested that the ancestors of the Khoi peoples are relatively recent pre-Bantu agricultural immigrants to Southern Africa, who abandoned agriculture as the climate dried and either joined the San as hunter-gatherers or retained pastoralism.
Since the arrival of the Bantu expansion in Southern Africa about 1,500 years ago, linguistic influence is seen in the adoption of click consonants and loan words from Khoisan into the Xhosa and Zulu languages. Bantu-speaking communities would have reached southern Africa from the Congo basin by about the 6th century AD. The advancing Bantu encroached on the Khoikhoi territory, forcing survivors of the indigenous populations to move to more arid areas of the Kalahari.
Their husbandry of sheep, goats and cattle grazing in fertile valleys across the region provided a stable, balanced diet, and allowed the Khoikhoi to live in larger groups in a region previously occupied by the San, who were subsistence hunter-gatherers. Advancing Bantu in the 3rd-6th century AD encroached on the Khoikhoi territory, pushing them into more arid areas.
The Bantu people, with advanced agriculture and metalworking technology, outcompeted and intermarried with the Khoisan, becoming the dominant population of South-Eastern Africa before the arrival of the Dutch colonists in 1652.
After the arrival of the Bantu, the Khoisan and their pastoral or hunter-gatherer ways of life remained predominant west of the Fish River in South Africa and in deserts throughout their region, where the drier climate precluded the growth of Bantu crops suited for warmer and wetter climates.
San woman in Namibia (1984 photograph)
The Khoikhoi enter the historical record with their first contact with Portuguese explorers,
about 1,000 years after their displacement by the Bantu. Local population dropped after the Khoi were exposed to smallpox from Europeans. The Khoi waged more frequent attacks against Europeans when the Dutch East India Company enclosed traditional grazing land for farms.
Khoikhoi social organisation was profoundly damaged and, in the end, destroyed by colonial expansion and land seizure from the late 17th century onwards. As social structures broke down, some Khoikhoi people settled on farms and became bondsmen (bondservants) or farm workers; others were incorporated into existing clan and family groups of the Xhosa people.
Georg Schmidt, a Moravian Brother from Herrnhut, Saxony, now Germany, founded Genadendal in 1738, which was the first mission station in southern Africa, among the Khoi people in Baviaanskloof in the Riviersonderend Mountains. Early European settlers sometimes intermarried with Khoikhoi women, resulting in a sizeable mixed-race population now known as the Griqua.
Andries Stockenström facilitated the creation of the "Kat River" Khoi settlement near the eastern frontier of the Cape Colony. The settlements thrived and expanded, and Kat River quickly became a large and successful region of the Cape that subsisted more or less autonomously. The people were predominantly Afrikaans-speaking Gonaqua Khoi, but the settlement also began to attract other Khoi, Xhosa and mixed-race groups of the Cape.
The so-called "Bushman wars"
were to a large extent the response of the San after their dispossession.
At the start of the 18th century, the Khoikhoi in the Western Cape lived in a co-operative state with the Dutch. By the end of the century the majority of the Khoisan operated as 'wage labourers', not that dissimilar to slaves. Geographically, the further away the labourer was from Cape Town, the more difficult it became to transport agricultural produce to the markets. The issuing of grazing licences north of the Berg River in what was then the Tulbagh Basin propelled colonial expansion in the area. This system of land relocation led to the Khoijhou losing their land and livestock as well as dramatic change in the social, economic and political development.
After the defeat of the Xhosa rebellion in 1853, the new Cape Government endeavoured to grant the Khoi political rights to avert future racial discontent. The government enacted the Cape franchise in 1853, which decreed that all male citizens meeting a low property test, regardless of colour, had the right to vote and to seek election in Parliament. This non-racial principle was later abolished by the apartheid Government.
In the Herero and Namaqua genocide in German South-West Africa, over 10,000 Nama are estimated to have been killed during 1904–1907.
The San of the Kalahari were described in Specimens of Bushman Folklore by Wilhelm H. I. Bleek and Lucy C. Lloyd (1911).
They were brought to the globalised world's attention in the 1950s by South African author Laurens van der Post in a six-part television documentary.
The Ancestral land conflict in Botswana concerns the Central Kalahari Game Reserve (CKGR), established in 1961 for wildlife, while the San were permitted to continue their hunter-gatherer lifestyle. In the 1990s, the government of Botswana began a policy of "relocating" CKGR residents outside the reserve. In 2002, the government cut off all services to CKGR residents. A legal battle began, and in 2006 the High Court of Botswana ruled that the residents had been forcibly and unconstitutionally removed. The policy of relocation continued, however, and in 2012 the San people (Basarwa) appealed to the United Nations to force the government to recognise their land and resource rights.
Following the end of Apartheid in 1994, the term "Khoisan" has gradually come to be used as a self-designation by South African Khoikhoi as representing the "first nations" of South Africa vis-a-vis the ruling Bantu majority. A conference on "Khoisan Identities and Cultural Heritage" was organised by the University of the Western Cape in 1997. and "Khoisan activism" has been reported in the South African media beginning in 2015.
The South African government allowed Khoisan families (up until 1998) to pursue land claims which existed prior to 1913. The South African Deputy Chief Land Claims Commissioner, Thami Mdontswa, has said that constitutional reform would be required to enable Khoisan people to pursue further claims to land from which their direct ancestors were removed prior to 9 June 1913.