Even-toed ungulate

Even-toed ungulates
Temporal range: 55–0 Ma Early EoceneHolocene
GiraffeAmerican bisonRed deerKiller whaleWild boarDromedaryThe Artiodactyla.jpg
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Even-toed ungulates. While cetaceans such as whales and dolphins are not normal artiodactyls, they belong to the clade; the Artiodactyla are sometimes termed the Cetartiodactyla to reflect this.[1]
Scientific classification
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Infraclass:
(unranked):
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Artiodactyla

Owen, 1848
Subclades

The even-toed ungulates (Artiodactyla, from Ancient Greek ἄρτιος (ártios), meaning 'even', and δάκτυλος (dáktylos), meaning 'finger/toe') are ungulates (hoofed animals) whose weight is borne equally by the third and fourth toes. By contrast, odd-toed ungulates, such as horses, bear their weight primarily on their third toes. The aquatic cetaceans (whales, dolphins, and porpoises) evolved from even-toed ungulates, so modern taxonomic classification sometimes combines the Artiodactyla and Cetacea into the Cetartiodactyla.

Excluding whales, the roughly 220 artiodactyl species include pigs, peccaries, hippopotamuses, camels, llamas, alpacas, mouse deer, deer, giraffes, antelopes, sheep, goats, and cattle, many of which are of great dietary, economic, and cultural importance to humans.

Evolution

The oldest fossils of even-toed ungulates date back to the early Eocene (about 53 million years ago). Since these findings almost simultaneously appeared in Europe, Asia, and North America, it is very difficult to accurately determine the origin of artiodactyls. The fossils are classified as belonging to the family Dichobunidae; their best-known and best-preserved member is Diacodexis.[2] These were small animals, some as small as a hare, with a slim build, lanky legs, and a long tail. Their hind legs were much longer than their front legs. The early to middle Eocene saw the emergence of the ancestors of most of today's mammals.[3]

Two large boar-like creatures graze.
Entelodonts were stocky animals with a large head, and were characterized by bony bumps on the lower jaw.

Two formerly widespread, but now extinct, families of even-toed ungulates were Enteledontidae and Anthracotheriidae. Entelodonts existed from the middle Eocene to the early Miocene in Eurasia and North America. They had a stocky body with short legs and a massive head, which was characterized by two humps on the lower jaw bone. Anthracotheres had a large, porcine (pig-like) build, with short legs and an elongated muzzle. This group appeared in the middle Eocene up until the Pliocene, and spread throughout Eurasia, Africa, and North America. Anthracothereres are thought to be the ancestors of hippos, and, likewise, probably led a similar aquatic lifestyle. Hippopotamuses appeared in the late Miocene and occupied Africa and Asia – they never got to the Americas.[3]

The camels (Tylopoda) were, during large parts of the Cenozoic, limited to North America; early forms like Cainotheriidae occupied Europe. Among the North American camels were groups like the stocky, short-legged Merycoidodontidae. They first appeared in the late Eocene and developed a great diversity of species in North America. Only in the late Miocene or early Pliocene did they migrate from North America into Eurasia. The North American varieties became extinct around 10,000 years ago.

Suina (including pigs) have been around since the Eocene. In the late Eocene or the Oligocene, two families stayed in Eurasia and Africa; the peccaries, which became extinct in the Old World, exist today only in the Americas.

A deer-like animal wanders through a clearing.
Sivatherium was a group of giraffes with deer-like forehead weapons.

South America was settled by even-toed ungulates only in the Pliocene, after the land bridge at the Isthmus of Panama formed some three million years ago. With only the peccaries, lamoids (or llamas), and various species of capreoline deer, South America has comparatively fewer artiodactyl families than other continents, except Australia, which has no native species.

Other Languages
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Alemannisch: Paarhufer
aragonés: Artiodactyla
asturianu: Artiodactyla
Avañe'ẽ: Pysãkõi
azərbaycanca: Cütdırnaqlılar
Bân-lâm-gú: Ngó͘-tê tōng-bu̍t
башҡортса: Ҡуш тояҡлылар
беларуская: Парнакапытныя
беларуская (тарашкевіца)‎: Парнакапытныя
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bosanski: Papkari
brezhoneg: Artiodaktiled
català: Artiodàctils
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čeština: Sudokopytníci
Deutsch: Paarhufer
eesti: Sõralised
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interlingua: Artiodactylos
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Кыргызча: Ача туяктуулар
Latina: Artiodactyla
latviešu: Pārnadži
Lëtzebuergesch: Artiodactyla
lietuvių: Porakanopiai
Ligure: Artiodactyla
Limburgs: Evehovege
Lingua Franca Nova: Artiodatilo
македонски: Парнокопитни
Bahasa Melayu: Artiodactyla
Nederlands: Evenhoevigen
日本語: ウシ目
norsk: Klovdyr
norsk nynorsk: Klauvdyr
Novial: Artiodactyla
occitan: Artiodactyla
oʻzbekcha/ўзбекча: Juft tuyoqlilar
پنجابی: پدھرے کھر
Перем Коми: Вожьясянгыжаэз
português: Artiodátilos
română: Artiodactile
Runa Simi: Iskay ruk'anayuq
sicilianu: Artiodactyla
Simple English: Artiodactyl
slovenčina: Párnokopytníky
slovenščina: Sodoprsti kopitarji
српски / srpski: Папкари
srpskohrvatski / српскохрватски: Papkari
Tagalog: Artiodactyla
українська: Парнокопиті
Vahcuengh: Youdaezmoeg
Tiếng Việt: Bộ Guốc chẵn
Winaray: Artiodactyla
吴语: 偶蹄目
粵語: 偶蹄目
Zeêuws: Evenoevihen
中文: 偶蹄目